Dactylolabis (Coenolabis) posthabita subsp. tothi Kolcsar 2026, ssp. nov.

Dactylolabis (Coenolabis) posthabita tothi Kolcsár ssp. nov. Figs 1 F, 1 G, 2 H, 2 I, 3 G – K Dactylolabis (Coenolabis) posthabita (Bergroth, 1888) in Kolcsár et al. 2021. Type material. Holotype: Hungary • ♂ (pinned); Potony; 45.93°N, 17.64°E (approx.; not stated on the label); 03 May 1977; leg. S. Tóth; HOLOTYPE Dactylolabis (Coenolabis) posthabita tothi Kolcsár, ssp. nov. [red label]; HNHM. Paratypes: Hungary • 6 ♂♂, 1 ♀ (pinned); same locality and date as holotype; HNHM • 3 ♂♂ (pinned); Darány, Barcsi-Ősborókás; 45.98°N, 17.563°E (approx.; not stated on the label); 03 May 1977; leg. S. Tóth; HNHM • 2 ♀♀ (pinned); Pécs, Bányatelep; 46.13°N, 18.33°E (approx.; not stated on the label); 05–07 June 1957; leg. F. Mihály; det. as Dactylolabis tergestina Egger, 1863 by Mannheims 1965; HNHM • 1 ♀ (pinned); Pécs, Bányatelep, gesztenyés felett; 46.13°N, 18.33°E (approx.; not stated on the label); 05–07 June 1957; leg. F. Mihály; det. as Dactylolabis tergestina Egger, 1863 by Mannheims 1965; HNHM • 1 ♂ (pinned); Pécs, Mecsek-hg., Jakob hg., tölgyes; 46.09°N, 18.14°E (approx.; not stated on the label); 05–07 June 1957; leg. F. Mihály; det. as Dactylolabis tergestina Egger, 1863 by Mannheims 1965; HNHM. Diagnosis. Fused median part of interbase situated at approx. half-length of aedeagal complex (at two-thirds length in p. posthabita). Ventral arm of interbase rectangular, with a narrow, pointed apex in caudal view (drop-shaped, with a triangular apex in p. posthabita); as wide as tip of aedeagus and bent almost perpendicularly at mid-length in lateral view (narrow, band-like, with an evenly curved posterior margin in p. posthabita). Description. General appearance and size as nominate subspecies in both sexes (Fig. 1). Male terminalia. In general (Fig. 2 H, I, 3 G, H), similar to the nominate subspecies, differing as follows: interbase (Fig. 3 I) with fused median part situated at approx. half-length of aedeagal complex; in caudal view (Fig. 3 K) ventral arm of interbase longer than in nominate subspecies, nearly rectangular, directed more ventrally at approx. 45 ° with dorsal apex narrow, pointed; in lateral view (Fig. 3 J), as wide as apex of aedeagus, posterior margin bent almost perpendicularly at mid-length. Sperm pump as wide as aedeagus in lateral view (Fig. 3 J). Female. General appearance as male. Female terminalia. In general, similar to the nominate subspecies, differing as follows: lateral arms of genital frame elongate, approx. twice as long as wide in caudal view (Fig. 4 G). Etymology. The subspecies name is dedicated to the memory of the recently deceased Sándor Tóth (1930–2024), a Hungarian entomologist, who collected the majority of the types. The name is a noun in genitive case and masculine form. Discussion. The male genitalia of D. posthabita are very similar to those of D. imeretica, suggesting a closer morphological affinity between these taxa than between D. imeretica and D. aberrans. However, according to Savchenko (1969), females of D. imeretica cannot be distinguished from those of D. aberrans and therefore possess short hypogynial valves. In contrast, females of both subspecies of D. posthabita have distinctly elongate hypogynial valves. The two subspecies mainly differ in the shape and position of the ventral arm of the interbase. While the interbase appears identical in the paratype of p. posthabita from Slovenia and the specimen from Serbia, the posterior margin of the hypandrium differs; in the type material it is rounded in dorsal view (Fig. 2 E, G), whereas in the Serbian specimens it is produced into a finger-like lobe, clearly visible in lateral view (Fig. 3 C), which is also characteristic of D. imeretica. The Hungarian p. tothi Kolcsár, ssp. nov., shows an intermediate condition, with the posterior margin slightly produced (Fig. 3 H). Other observed differences visible in the figures, such as the shape of the proctiger, appear to result largely from deformation or slight differences in orientation. The wing venation of the examined specimens of both subspecies shows variation in the position of m – cu relative to the fork of M (Fig. 1 B, D, G). However, this character is highly variable among specimens and may even vary between the left and right wings of the same individual, suggesting that it is not associated with subspecific or population-level differences. The Hungarian specimens were collected in Potony, most likely along the Drava River or in a relatively natural forest patch in 1977 (Sándor Tóth pers. comm. 2018), with additional specimens collected a few kilometres further in an old natural Juniperus - dominated, open, sandy steppe nature reserve. Further specimens were collected in the Mecsek Mountains in oak and sweet chestnut forests. The Serbian specimens were collected near a small graveyard with large oak trees in the surrounding area; one male was observed resting on mosses at the base of a concrete fence. Both the Hungarian p. tothi Kolcsár, ssp. nov., and Serbian p. posthabita specimens were collected from early May to early June. The exact collection date of the Slovenian type material of p. posthabita is unknown; only the year is available. Erhan-Dincă (1984) reported that the Romanian specimens were collected in a wet, well-forested montane habitat between mid-September and late October. If the hypogynial valves of these specimens are indeed short, as stated by Savchenko (1986), then, considering both their occurrence at higher elevations and their markedly different flight period (autumn rather than late spring to early summer, as observed in other populations), the Romanian population could represent a distinct species and warrants further investigation.

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