Evidence for a Polyphyletic Origin of the Labiatae

AAnnals of the Missouri Botanical Garden Table 1.Taxa included in study group.Numbers in gynoecial structure occurs within Microcorys parentheses: number of genera included in this analysis/ (Prostanthereae).In the Verbenaceae, the ovary number of genera in the taxon.Parenthetical acronyms is usually unlobed but may be lobed as much as are used in Table 2 andFigures 1-4.halfway to the base (e.g., some species of Oxera).-In summary, there is a continuum in the degree of ovary lobing, with some intermediates assigned to the Labiatae and some to the Verbenaceae.Since this is the orJy character distinguishing the two famihes as currently circumscribed, the taxonomic limits of the Labiatae are unclear, and there is no Verbenaceae sensu lato Caryopteridoideae (6/6) Caryopterideae (5/5) (CAR) Teijsmanniodendreae (1/1) (TEI) Chloanthoideae (10/10) Chloantheae (5/5) (CH) Physopsideae (5/5) (PH) Verbenoideae (2/ca.38) MonochUeae (2/2) (MO) Viticoideae (26/28) Callicarpeae (3/4) (CAL) Clerodendreae (10/10) (CL) Tectoneae (3/3) (TEC) Viticeae (10/11) (VI) Labiatae Lamioideae (32/ca.83) Ajugeae (12/12) (AJ) Prostanthereae (7/7) (PR) Scutellarieae (4/4) (SC) Gynobasic-styled Lamioideae (8/ca.60) (GL) Nepetoideae (4/ca.160) (NE)

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